Where $`Flux_{i,j}`$ is the increase in concentration in compartment $`i`$ [mmol-N m<sup>-3</sup> d<sup>-1</sup>] due to the water exchange between compartment $`i`$ and $`j`$. $`Ex_{i,j}`$ is the exchange rate between compartment $`i`$ and $`j`$ [m<sup>3</sup> d<sup>-1</sup>]. $`C_{i}`$ and $`C_{j}`$ are the concentrations of the state variables in compartment $`i`$ and $`j`$ [mmol-N m<sup>-3</sup>], respectively. $`Vol_{i}`$ is the volume of compartment $`i`$ [m<sup>3</sup>].
Where $`FluxC_{i,j}`$ is the increase in concentration of state variable $`C`$ in compartment $`i`$ [mmol-N m<sup>-3</sup> d<sup>-1</sup>] due to the water exchange between compartment $`i`$ and $`j`$. $`Ex_{i,j}`$ is the exchange rate of water between compartment $`i`$ and $`j`$ [m<sup>3</sup> d<sup>-1</sup>]. $`C_{i}`$ and $`C_{j}`$ are the concentrations of state variable $`C`$ in compartment $`i`$ and $`j`$ [mmol-N m<sup>-3</sup>], respectively. $`Vol_{i}`$ is the volume of compartment $`i`$ [m<sup>3</sup>].