DINupMPB is calculated from the maximum specific uptake rate ($`MPB_{upt,max}`$) [d<sup>-1</sup>] at ambient temperature and the algal concentration. Limitations by light and nutrients are described by Monod formulations.
Where $`MPB_{upt,max} `$ is the maximum DIN uptake rate [d<sup>-1</sup>] of microphytobenthos at 10°C, $`ks_{PAR,MPB}`$ and $`ks_{DIN,MPB}`$ are the half-saturation coefficients for light and DIN, respectively. Production of the microphytobenthos is limited by the carrying capacity ($`CC_{MPB}`$) [mmol N m<sup>-2</sup>] at the intertidal flat. $`T_{fac}`$ is the correction for the ambient water temperature.